Galapagos Species Database
The Galapagos Species Database shares the information about the species from our Natural History Collections.
Howardia biclavis
Escama, Mining scale
Domain
Eukaryota
Kingdom
Animalia
Phylum
Arthropoda
Class
Insecta
Order
Hemiptera
Suborder
Sternorrhyncha
Superfamily
Coccoidea
Family
Diaspididae
Genus
Howardia
Species
biclavis
Taxon category: Accepted
Origin: Introduced - established
Year of first record: 1975
Mode of introduction: Accidental
Introduction Pathway: Contaminant
Subpathway: Contaminant on plants (inc. seeds and plant associated material)
Introduced status: Naturalized
Invasive status: Lesser concern
Substrate or host preferences: The Galapagos host plants: Vachellia macracantha (Acacia), Chiococca sp., Citrus sinensis (Orange x Persian lime hybrid), Croton sp., Waltheria ovata.
Map of specimen collection localities or observation records for this species in our collections database.
Distribution: Santiago, San Cristobal, Santa Cruz
- Lincango, M.P. Hodgson, C. & Causton, C. (2008) Lista anotada de los insectos escama (Hemiptera: Coccoidea) de las islas Galápagos, Ecuador. Charles Darwin Research Station, unpublished, 17 pp.
- Lincango, P. Hodgson, C., Causton, C. & Miller, D. (2010) An updated checklist of scale insects (Hemiptera: Coccoidea) of the Galapagos Islands, Ecuador. Galapagos Research, 67: 3-7.
- Peck, S.B. (1996) Origin and development of an insect fauna on a remote archipelago: The Galápagos Islands, Ecuador. In: Keast A., Miller S.E. (eds.): The origin and evolution of Pacific Island biotas, New Guinea to eastern Polynesia: patterns and processes. SPB Academic Publishing, Amsterdam, The Netherlands, p. 91-122.
- Peck, S.B. Heraty, J., Landry, B. & Sinclair, B.J. (1998) Introduced insect fauna of an oceanic archipelago: The Galápagos Islands, Ecuador. Am. Entomol. 44: 218-237.
- Peck, S.B. (2001) Smaller Orders of Insects of the Galápagos Islands, Ecuador: Evolution, Ecology, and Diversity. NRC Research Press, Ottawa, Ontario, Canada, 278 pp.
- Williams, M.L. (1977) Scale insects of the Galapagos islands In D.R Miller, J.A Davidson, M. Kosztarab (Eds.), Morphology and Systematics of scales insects (Ed.127, Vol. 9, pp. 85-98). Research Division Bulletin, Virginia Polytechnic Institute, and State University.
You are welcome to download and use the information found in this page, acknowledging its source.
This page should be cited as follows:
"Galapagos Species Database, Howardia biclavis", dataZone. Charles Darwin Foundation, https://datazone.darwinfoundation.org/en/checklist/?species=11102. Accessed 31 October 2026.