Galapagos Species Database

The Galapagos Species Database shares the information about the species from our Natural History Collections.

Ramalina darwiniana var. darwiniana Aptroot & Bungartz

ramalina, Ramalina

Thallus shrubby, greenish grey, decaying parts becoming red, without a distinct holdfast, relatively densely branched, up to 15 cm diam., but usually much smaller (3-7 cm); branches initially upright, hanging down when older, in most parts distinctly canaliculate and in section broadly crescent-shaped to±flattened, sometimes also markedly curved in the longitudinal direction and frequently with curved lobe tips (“shepherds' crooks”), slender to relatively broad (up to 15 mm wide), occasionally anastomosing, leaving oval to elongated holes; lobe margins with rounded, irregular warts from which irregular, punctiform or sublinear pseudocyphellae develop; soralia absent. Apothecia common, c. 2–7 mm diam., cupular, laminal to subterminal; disc yellowish to pinkish grey; branches often bent at the apothecia (geniculate), verrucose below with crateriform pseudocyphellae; ascospores hyaline, 1-septate, narrowly oblong, usually ±curved, 10-12 x 4–5 µm. Pycnidia mostly marginal, hemispherical, 0.1–0.2 mm diam., ostiole pale ochre; conidia rod-shaped, 5–6 x 0.5–1.0 µm. Chemistry: Medulla K+ orange yellow, soon red, UV+ white or UV- (salazinic, sekikaic, divaricatic acid); cortex with usnic acid, occasionally (e.g. in the type collection) also with atranorin.

The species is similar to R. complanata, which, however, has an irregularly flattened, not distinctly canaliculate surface and laminal tuberculate pseudocyphellae. Ramalina sideriza is also vaguely similar in habitus and as with R. darwiniana has hooked lobe tips, but it has flattened lobes with a distinctly striate, often coarsely linear pseudocyphellate surface.

Taxonomy

Domain
Eukaryota

Kingdom
Fungi

Phylum
Ascomycota

Class
Lecanoromycetes

Order
Lecanorales

Family
Ramalinaceae

Genus
Ramalina

Species
darwiniana

Subspecies
var. darwiniana

Taxon category: Species with Infraspecific Taxa

Type: ECUADOR, Galapagos, Plaza Sur Island, 0º34’59”S, 90º9’54”W, alt. 1 m, coastal zone; eastern part with scattered and low vegetation of Sesuvium portulacastrum & Tiquilia galapagoa with occasional Opuntia, western part also with scattered shrubs of Grabowskia boerhaaviaefolia, Maytenus octogona, and Castela galapageia, on wood, twig, 21-Feb-2006, A. Aptroot, 64433 (CDS no. 31001—holotype; ABL—isotype).

Origin: Endemic

Status

Least concern

Distribution

Map of specimen collection localities or observation records for this species in our collections database.

Distribution: R. darwiniana is corticolous or lignicolous, rarely also saxicolous, and grows throughout the coastal and arid zone, rarely higher. It is often abundant on twigs and branches in coastal shrub and dry forest.

References

  • Aptroot, A. Bungartz, F. (2007) The Lichen Genus Ramalina on the Galapagos. The Lichenologist 39: 519-542.

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This page should be cited as follows:

"Galapagos Species Database, Ramalina darwiniana var. darwiniana", dataZone. Charles Darwin Foundation, https://datazone.darwinfoundation.org/en/checklist/?species=3318. Accessed 11 August 2025.