Galapagos Species Database
The Galapagos Species Database shares the information about the species from our Natural History Collections.
Phaeographis atromaculata
liquen de escritura, Script lichen

Thallus thin, often partially embedded in the bark, continuous to rimose, pale creamy white, grayish or ochraceous, ± the same color as the substrate; surface smooth, matt, not pruinose; Apothecia immersed to prominent, lirellate; lirellae broad, short to ±elongate, sparsely branched; labia absent, but with a smooth, prominent thalline margin; disc expanded, flattened, blackened, typically ±covered by an inconspicuous farinose whitish pruina; exciple thin, laterally and basally not carbonized, not expanding, but apically pigmented, ±brownish, not black, typically surrounded by a smooth thalline margin; hymenium hyaline, strongly inspers, I–; ascospores pale gray, soon brown, I+ violet or brownish violet, 1-4 per ascus, broadly oblong, 95-128 x 20-32 µm, densely muriform, 21-31 vs. 4-7 celled. Chemistry: Thallus with norstictic and connorstictic acid, K+ orange-red (crystals).
Because of its broadly expanded, inconspicuously pruinose disc and sparsely branched lirellae P. atromaculata is superficially similar to Sarcographa ramificans. Labia of P. atromaculata are, however, not well developed, instead an often fissured whitish thallus rim typically surrounds the disc. The two species are also clearly distinguished by their secondary chemistry (S. ramificans lacks norstictic acid). Microscopically, P. atromaculata has large, densely muriform ascospores, the exciple is not carbonized and basally poorly developed. Sarcographa ramificans has much smaller, transversely septate spores and a distinctly carbonized exciple that basally expands with age.
Domain
Eukaryota
Kingdom
Fungi
Phylum
Ascomycota
Class
Lecanoromycetes
Order
Ostropales
Family
Graphidaceae
Genus
Phaeographis
Species
atromaculata
Taxon category: Accepted
Syn.: Phaeographis kalbii, fide Bungartz et al. (2009)
Origin: Native
Map of specimen collection localities or observation records for this species in our collections database.
Distribution: A pantropical species; typical for the transition zone, on Bursera graveolens, Opuntia galapageia ssp. macrocarpa, Trema micrantha, Chiococca alba, and Croton scouleri.
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This page should be cited as follows:
"Galapagos Species Database, Phaeographis atromaculata", dataZone. Charles Darwin Foundation, https://datazone.darwinfoundation.org/en/checklist/?species=3956. Accessed 5 July 2025.